User blog comment:Thailog/May episodes details/@comment-1185312-20120428005725/@comment-1635895-20120428023753

Greg confirmed it in an early interview.

"Aqualad came from the Atlantean city of Shayeris, where he was raised by Sha'lain'a, a native of that city, and Calvin Durham, a henchman of Black Manta who was genetically altered to infiltrate Atlantis. However, his actual father was Black Manta himself." Source.